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Carlos M Gradil - LMV, MS, PhD, DACT

Carlos M Gradil
Reproductive Consultant in Theriogenology/Reproduction

Campus Phone:



  • PhD in Theriogenology - University of Minnesota - 1987
  • MSc in Theriogenology - University of Minnesota - 1982
  • LMV - University of Lisbon, Portugal - 1977

Board Certification

  • Diplomate of the American College of Theriogenology - 1998
  1. Gradil, C.M., Uricchio, C.K., Schwarz, A. 2019. Self-Assembling Intrauterine Device (Upod) Modulation of the Reproductive Cycle in Mares. Journal of Equine Veterinary Science.
  2. Frank, N., Hermida, P., Sanchez-Londoño, A., Singh, R., Gradil, C.M., Uricchio, C.K. 2017. Blood Glucose and Insulin Concentrations after Octreotide Administration in Horses With Insulin Dysregulation. Journal of Veterinary Internal Medicine.
  3. Pirie, C.G., Alario, A.F., Barysauskas, C.M., Gradil, C., Uricchio, C.K. 2014. Manual corneal thickness measurements of healthy equine eyes using a portable spectral-domain optical coherence tomography device. Equine Veterinary Journal.
  4. Gradil, C., McCarthy, R. 2013. Cryptorchidism. Small Animal Soft Tissue Surgery.
  5. O'Connor, C.L., Schweizer, C., Gradil, C., Schlafer, D., Lopate, C., Prociuk, U., Meyers-Wallen, V.N., Casal, M.L. 2011. Trisomy-X with estrous cycle anomalies in two female dogs. Theriogenology.
  6. Coyne, M.J., Cousin, H., Loftus, J.P., Johnson, P.J., Belknap, J.K., Gradil, C.M., Black, S.J., Alfandari, D. 2009. Cloning and expression of ADAM-related metalloproteases in equine laminitis. Veterinary Immunology and Immunopathology.
  7. Frederico, L.M., Gerard, M.P., Pinto, C.R.F., Gradil, C.M. 2007. Bilateral occurrence of granulosa-theca cell tumors in an Arabian mare. Canadian Veterinary Journal.
  8. Volkmann, D.H., Gilbert, R.O., Gradil, C., Linke, R. 2002. Leukocytes trapped in the fetal membranes of newborn foals. Theriogenology.
  9. Gradil, C.M., Ball, B.A. 2000. The use of pentoxifylline to improve motility of cryopreserved equine spermatozoa. Theriogenology.
  10. Gradil, C.M., Watson, R.E., Renshaw, R.W., Gilbert, R.O., Dubovi, E.J. 1999. Detection of bovine immunodeficiency virus DNA in the blood and semen of experimentally infected bulls. Veterinary Microbiology.
  11. Gradil, C., Dubuc, C., Eaglesome, M.D. 1996. Porcine reproductive and respiratory syndrome virus: Seminal transmission. Veterinary Record.
  12. Harding, M.J., Prud'homme, I., Gradil, C.M., Heckert, R.A., Riva, J., McLaurin, R., Dulac, G.C., Vydelingum, S. 1996. Evaluation of nucleic acid amplification methods for the detection of hog cholera virus. Journal of Veterinary Diagnostic Investigation.
  13. Gradil, C., Eaglesome, M.D., Stewart, B., Garcia, M.M., Quimby, F. 1995. Bactericidal effects of ozone at nonspermicidal concentrations.. Canadian journal of veterinary research = Revue canadienne de recherche vétérinaire.
  14. Gradil, C.M., Harding, M.J., Lewis, K. 1994. Use of polymerase chain reaction to detect porcine parvovirus associated with swine embryos.. American journal of veterinary research.
  15. Gradil, C., Sampath, M., Eaglesome, M.D. 1994. Detection of verotoxigenic Escherichia coli in bull semen using the polymerase chain reaction. Veterinary Microbiology.
  16. Bielanski, A., Sampath, M., Gradil, C., Eaglesome, M.D., Garcia, M. 1994. In vitro Fertilization of Bovine Ova in the Presence of Campylobacter fetus subsp. venerealis. Reproduction in Domestic Animals.
  17. Gradil, C., Molitor, T., Harding, M., Crabo, B. 1990. Excretion of porcine parvovirus through the genital tract of boars.. American journal of veterinary research.
  18. Gradil, C.M., Joo, H.S., Molitor, T.W. 1990. Persistence of Porcine Parvovirus in Swine Infected In Utero and Followed through Maturity. Journal of Veterinary Medicine, Series B.
  19. Bane, D.P., James, J.E., Gradil, C.M., Molitor, T.W. 1990. In vitro exposure of preimplantation porcine embryos to porcine parvovirus. Theriogenology.
  20. Gradil, C., Joo, H.S. 1988. A radial immunodiffusion enzyme assay for detection of antibody to equine rhinopneumonitis virus (EHV-1) in horse serum. Veterinary Microbiology.

Research and Clinical Interests

  • Breeding management, artificial insemination with fresh, chilled and frozen semen.
  • Transcervical insemination.
  • Breeding soundness evaluation.
  • Collection and processing of semen for shipment.
  • Semen cryopreservation (freezing).
  • Pregnancy diagnosis.